We will train 15 Early Stage Researchers (ESR) in these skills, with a training plan to give them the required expertise in the relevant areas, and their combination. We will pursue four research themes, two developing methodology and two focused on applications.

The first concerns the modelling of structured populations, incorporating more realistic spatial and social interactions. The second considers important out of equilibrium dynamical concepts, often neglected in favour of equilibria. The third focuses on the mathematical modelling of cancer and its treatment. Here the two above concepts are especially important. The fourth considers ecological and epidemiological modelling where the focus is on structural complexity relating to the interplay of different timescales and the management and analysis of epidemiological data. Whilst specialising, all ESRs will gain knowledge of all of these important themes.